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The single player is modeled after the Normandy breakout, where the British, Canadian, Polish, and American forces pushed into the city of Chambois, France, also known as the Falaise Gap. Unlike most other games in the Call of Duty series, the events in Call of Duty 3 are based on a single combined campaign, with the player being switched between the four nations and their respective characters for each leg of the story. There are 14 playable campaign missions.
In the American missions, the player assumes control of Private Nichols, recently arrived in France and eventually attached to the 29th Infantry Division. Nichols and his squad participate in the capture of Saint Lo. After that the squad is transferred to the 90th Infantry Division and sent to secure the wooded area of Saint Germain-Sur-Seves, where intense hedgerow fighting took place. Later, the unit is transferred to the town of Mayenne, where team mate Pvt. Huxley is tasked with defusing bombs planted on several bridges. Huxley is wounded before he can carry out the order, and Sergeant McCullin, the unit leader and also a former combat engineer, volunteers to defuse the bombs. McCullin successfully defuses the bombs, but is mortally wounded; Corporal Dixon assumes command of the unit. The squad is later tasked with clearing out a forest and finding the engineers sent in before that have not been heard from. The squad participates in going to a nearby town to clear it out, Dixon is wounded here, but recovers. The unit is then sent to defend the town of Chambois in a brutal battle. Private First Class Guzzo, another member of the unit, performs the suicidal task of laying colored smoke grenades on German positions while under heavy fire in order to direct air support. He is wounded, but is carried to safety by Dixon. While treating Guzzo, Dixon is shot. He passes on the final words of McCullin, as well as his own, to Guzzo, who is in charge of the unit and its replacements by the end of the game.