The X360's XBL is the only system where one can has an always on external, independent chat system and the speculation is MS wants to do the same thing for externals aps, which would of course including non-gaming stuff, but could also include video chat, game screenshots, game recording and even streaming of games.
http://www.gamesindustry.biz/articles/digitalfoundry-the-os-factor
Also, expect both Sony and Microsoft to bolster the ability to share in-game video and screenshot assets in their next-gen hardware. Most high-end GPUs these days have hardware video encoders built-in and this functionality is a natural fit for next-gen consoles: players may even be able to livestream their own gameplay. An onboard video encoder would have useful, if rather more mundane uses too: the same tech could be used to re-encode downloaded video to beam across to other devices for example, and even power videochat during gameplay (don't forget that MS now owns Skype). On a more practical level, the ability to merge OS-level programs and gameplay could allow for strategy guides to become far more contextual in nature, offering up in-game maps and video based on your actual level of progress.
As a point of reference, if true the X720 would have 6x the game RAM of the Wii U and twice the system RAM.
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