MGS2 followed closely by MGS3.
Gameplay wise MGS2 is a less ambitious game than MGS3, but it's a tighter and more refined experience all around.
I also much prefer the story in MGS2, but both games do have very enjoyable stories for very different reasons.
As others have mentioned, GZ and the upcoming TPP take the gameplay king spot.
MGS1 will always be the classic, but it doesn't stand toe to toe with these games on any level. MGS2 takes the gameplay style of MGS1, but expands and perfects it. MGS3, while less refined than 2, still glides right over the original with its breadth of mechanics and relatively open ended level design. Some argue that the story in MGS1 is the most down to earth, but it's still really damn stupid when you realize that the Foxhound members are sacrificing themselves and Liquids plan is to let you win up until REX is activated.
Peace Walker? Ridiculous story, but a fun game. Neutered MGS3 with a ton of content and some fun rpg elements.
MGS4? Garbage story. Good mechanics that rarely get put to good use due to 1. Abysmal level design and 2. Game breaking Octocamo system. Lots of great ideas with terrible implementation, you could tell Kojima was only able to get a shadow of his original design ideas into the final game. The open ended, ever changing, always aggressive battle field gameplay never quite panned out. Kind of sucks because it seems his game plan for MGS4 would have worked perfectly with the Fox Engine.
Portable Ops = pure shit all around. A significant era of the timeline wasted on a garbage game.
Metal Gear blows, don't care if it's the original.
Metal Gear 2: Solid Snake is actually a good game. Might as well be MGS1 in 2D. It's the Link to the Past to Ocarina. The Super Metroid to Metroid Prime.
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